Atlas of Breeding Birds of the
Sierra de Guadarrama National Park

III ATLAS OF BIRDS
during the breeding season in Spain

 

The fieldwork was carried out across the 21,714 hectares occupied by the Sierra de Guadarrama National Park within the Community of Madrid. Of the 330 grid squares that include part of their surface inside the park, surveys were conducted in 282 of them in order to determine the complete species list and the relative abundance of birds in each one. The 48 grid squares that were not visited correspond to those with a very small area inside the National Park. Within each surveyed square, all environments or habitats present were explored, since each one may host different species.

This national park lies at a considerable altitude above sea level (900–2,490 m), which strongly influences the types of habitat found there. Alpine grasslands, humid meadows, and mountain grasslands predominate, but there are also large forested areas. Pine and oak forests are common, and holm oak stands appear to a lesser extent. Shrubland is also an important component, especially dominated by broom and heath.

 

A total of 98 species were recorded, detailed in Annex 1. All of them are breeding species in the area. The distribution map and the habitats used by each species are included in the section Species Index.

The most widely distributed or most common species, according to the number of grid squares and in order of importance, are: common chaffinch, coal tit, European robin, rock bunting, dunnock, Eurasian wren, crested tit, and Eurasian blackcap. These eight species are present in more than 35% of the grid squares within the study area. In addition to these, three major groups can be identified:

  • More or less common species (21), present in 15% to 50% of the grid squares (Annex 1): rock bunting, dunnock, Eurasian wren, crested tit, Eurasian blackcap, short-toed treecreeper, firecrest, woodlark, griffon vulture, mistle thrush, common blackbird, skylark, European stonechat, western Bonelli’s warbler, common wood pigeon, Eurasian jay, common cuckoo, cinereous vulture, northern wheatear, common raven, and great tit.
  • Uncommon species (19), present in only 5% to 15% of the grid squares (Annex 1): great spotted woodpecker, goldcrest, common linnet, ortolan bunting, carrion crow, black redstart, Eurasian nuthatch, red crossbill, Eurasian blue tit, European serin, western subalpine warbler, long-tailed tit, western Orphean warbler, tawny pipit, common chiffchaff, common whitethroat, booted eagle, Dartford warbler, and common rock thrush.
  • Very localised species (55), present in less than 5% of the grid squares (Annex 1): water pipit, common nightingale, garden warbler, pied flycatcher, red-legged partridge, Iberian green woodpecker, red-billed chough, citril finch, common buzzard, blue rock thrush, common swift, red kite, bluethroat, common quail, crag martin, tree pipit, Sardinian warbler, melodious warbler, grey wagtail, scrub warbler, cirl bunting, red-backed shrike, Iberian magpie, song thrush, European goldfinch, spotless starling, golden eagle, black kite, European nightjar, red-rumped swallow, northern goshawk, Eurasian sparrowhawk, corn bunting, European greenfinch, rock sparrow, house sparrow, common kestrel, white wagtail, great grey shrike, Eurasian hoopoe, short-toed snake eagle, Thekla’s lark, barn swallow, western yellow wagtail, Eurasian golden oriole, peregrine falcon, European honey-buzzard, Bonelli’s eagle, hawfinch, woodchat shrike, spotted flycatcher, Spanish imperial eagle, white-throated dipper, Eurasian magpie, and European turtle dove.

In addition to the species recorded with signs of breeding, other species were also identified that are considered migratory or incidental observations, given the mobility of this group of fauna, which can result in sightings of dispersed individuals of almost any species anywhere. These include: common house martin, common redstart, willow warbler, greater short-toed lark, Eurasian collared dove, and brambling.

Species inventory by UTM 1×1 km grid

The number of species present in each grid square represents the specific richness per area in the park, or the species richness of each individual square. The 1×1 km grid squares shown in dark colours in Figure 1 represent the areas with the highest species richness in the national park.

The grid squares with the highest number of species (more than 20 recorded species) are distributed across different zones of the park, following patches of vegetation with greater diversity and more varied environments. Areas with lower species richness correspond to higher altitudes with little or no vegetation. Notable areas with higher diversity include the oak forests on the slopes extending above the village of Rascafría and the pine forests on both sides of the Navafría pass. Higher-richness areas also occur on the Morcuera slope towards Rascafría, the area known as El Pinganillo, and especially Las Hoyuelas (Figures 1, 2, and 3). Equally notable are the species-rich areas of the Hueco de San Blas and the southern slopes of Morcuera. The species found in these forested areas constitute typical mountain forest communities, including birds native to the predominant pine forest, as well as species from mixed forests, oak woods, and shrublands where these formations occur.

Most of the grid squares with a low number of species (1 to 5), and a large proportion of those with few species (6 to 10) are located mainly in the highest areas, that is, the peaks of the National Park, and in rocky slopes with little vegetation. These areas correspond to zones dominated by rocky habitats, high mountain grasslands, and broom scrub. In these habitats, species richness and bird abundance decline markedly, although broom scrub areas can host high densities of birds during the breeding season, with skylarks, dunnocks, and European stonechats being the main representatives.

A group of species consistently present within the park boundaries are the birds of prey. The colonies of cinereous vulture in Rascafría and La Pedriza, along with the small colonies of griffon vulture distributed across various rocky areas of the mountains, contribute a significant number of individuals, and their ease of detection means that visitors are almost constantly observing large birds in the park. Other birds of prey in higher threat categories, such as the Spanish imperial eagle or Bonelli’s eagle, are represented much more sparsely.

The presence of rare and striking species such as the common rock thrush and the bluethroat is also notable. Some species that used to be more abundant, although detected during this atlas’ fieldwork, are now in decline and scarcely represented; this is the case of the alpine accentor.

 
Figure 1. Species richness per 1×1 km UTM grid.
 
 
Figure 2. Species richness per 1×1 km UTM grid.
 
 
Figure 3. Names used in the text for some areas of the national park.

 

Threatened species and their distribution

Las especies de fauna están catalogadas de distinta manera en los diferentes catálogos de especies protegidas según la escala (estatal o regional) o según criterios específicos estatales o internacionales (Acuerdo del Consejo de Ministros de 24 de febrero de 2017, por el que se aprueban los criterios orientadores para la inclusión de taxones y poblaciones en el Catálogo Español de Especies Amenazadas o UICN).

Para que quede debidamente reflejada la distribución de las especies según estos criterios, a continuación se incluyen los listados y la distribución de las especies según su grado de amenaza considerando los siguientes documentos: Catálogo Español de Especies Amenazadas y Listado de Especies Silvestres en Régimen de Protección Especial, Libro Rojo de las Aves de España, Catálogo Regional de especies amenazadas de fauna y flora silvestres de la Comunidad de Madrid.

Threatened species according to the Spanish Catalogue of Threatened Species

Of the 98 species present in the Park during the breeding season, 78 are included in the List of Wild Species under Special Protection, and 4 of them appear in the Spanish Catalogue of Threatened Species with a very unfavourable conservation status: Endangered (2 species) and Vulnerable (2 species; tables 1, 2 and 3). The full list of species and their category is included in Annex 2.

Category Number of species
Endangered 2
Vulnerable 2
Included in the List of Wild Species under Special Protection 78

Table 1. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the Spanish Catalogue of Threatened Species in Spain.

Endangered

Spanish imperial eagle

Red kite

Table 2. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Spanish Catalogue of Threatened Species in Spain.

Vulnerable

Cinereous vulture

Bonelli’s eagle

Table 3. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Vulnerable according to the Spanish Catalogue of Threatened Species in Spain.

 

Considering the Spanish Catalogue of Threatened Species, the following areas would be identified as occupied by species in the highest threat categories within that catalogue:

Figure 4. Areas with the presence of species classified as Endangered in the Spanish Catalogue of Threatened Species.

Figure 5. Areas with the presence of species included in the Vulnerable category in the Spanish Catalogue of Threatened Species.

 

Threatened species according to the Red Book of the Birds of Spain

Of the 98 species present in the National Park during the breeding season, 14 are in an unfavourable conservation status and are included in the threat categories of Endangered (7 species) and Vulnerable (7 species) in the latest Red List prepared by SEO/BirdLife for the Ministry for the Ecological Transition and the Demographic Challenge (tables 4, 5 and 6).

Category Number of species
Endangered 7
Vulnerable 7
Near Threatened 13
Least Concern 70
Data Deficient 2

Table 4. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).

Endangered

Spanish imperial eagle

Woodchat shrike

Great grey shrike

Common kestrel

Common quail

Dartford warbler

Red kite

Table 5. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).

Vulnerable

Bonelli’s eagle

Red-backed shrike

Skylark

Barn swallow

Red-legged partridge

European turtle dove

Common swift

Table 6. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Vulnerable in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).

 

The distribution of species included in any unfavourable conservation status must guide the management of a protected area. In this case, the species detected as breeders within the boundaries considered in this study, and included in two of the highest categories according to IUCN criteria, highlight the priority conservation areas in this regard (figures 6 and 7).

Figure 6. Grid squares with the presence of species classified as Endangered in the latest Red Book.

Figure 7. Grid squares with the presence of species included in the Vulnerable category in the latest Red Book.

 

Threatened species according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid

Of the 98 species present in the National Park during the breeding season, 17 are in an unfavourable conservation status and are included in the threat categories of Endangered (3 species), Vulnerable (2 species), Special Interest (1 species), and Sensitive to Habitat Alteration (11 species) in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid (tables 7 and 8).

Category Number of species
Endangered 3
Vulnerable 2
Special Interest 1
Sensitive to Habitat Alteration 11

Table 7. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.

Endangered

Spanish imperial eagle

Bonelli’s eagle

Cinereous vulture

Table 8. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.

 

Considering the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid, the following areas would be identified as occupied by species in the highest categories of that catalogue:

Figure 8. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.

Figure 9. Grid squares with the presence of species included in the Vulnerable category in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.

Spanish Catalogue of Threatened Species and List of Wild Species under Special Protection:

EN – Endangered, VU – Vulnerable, LSPE – Included in the List of Wild Species under Special Protection.

Red Book of the Birds of Spain:

CR – Critically Endangered, EN – Endangered, VU – Vulnerable, DD – Data Deficient, NT – Near Threatened

Regional Catalogue of Threatened Species of Wild Fauna and Flora:

EN – Endangered, SAH – Sensitive to Habitat Alteration, VU – Vulnerable, DIE – Special Interest.

Annex 1

List of bird species present in the National Park. It is indexed by species name for quick identification and consultation of a specific species.

Scientific name Common name (Spanish) CEA 2024 Madrid 1992 Red Book 2021
Accipiter gentilis Northern goshawk LSPE   LC
Accipiter nisus Eurasian sparrowhawk LSPE   LC
Aegithalos caudatus Long-tailed tit LSPE   LC
Aegypius monachus Cinereous vulture VU Endangered NT
Alauda arvensis Skylark LSPE   VU
Alectoris rufa Red-legged partridge LSPE   VU
Anthus campestris Tawny pipit LSPE   VU
Anthus spinoletta Water pipit LSPE   LC
Anthus trivialis Tree pipit LSPE   LC
Apus apus Common swift LSPE   LC
Aquila adalberti Spanish imperial eagle EN Endangered VU
Aquila chrysaetos Golden eagle LSPE Sensitive to Habitat Alteration LC
Aquila fasciata Bonelli’s eagle VU Endangered VU
Buteo buteo Common buzzard LSPE   LC
Caprimulgus europaeus European nightjar LSPE   LC
Carduelis carduelis European goldfinch LSPE   LC
Carduelis citrinella Citril finch LSPE Special Interest NT
Cecropis daurica Red-rumped swallow LSPE   LC
Certhia brachydactyla Short-toed treecreeper LSPE   LC
Chloris chloris European greenfinch LSPE   LC
Cinclus cinclus White-throated dipper LSPE Special Interest LC
Circataetus gallicus Short-toed snake eagle LSPE   LC
Cocothraustes cocothraustes Hawfinch LSPE Special Interest LC
Columba palumbus Common wood pigeon LSPE   LC
Corvus corax Common raven     LC
Corvus corone Carrion crow     LC
Coturnix coturnix Common quail LSPE   LC
Cuculus canorus Common cuckoo LSPE   LC
Curruca communis Common whitethroat LSPE   LC
Curruca conspicillata Spectacled warbler LSPE   LC
Curruca hortensis Western Orphean warbler LSPE Special Interest LC
Curruca iberiae Western subalpine warbler LSPE   LC
Curruca melanocephala Sardinian warbler LSPE   LC
Curruca undata Dartford warbler LSPE   VU
Cyanistes caeruleus Eurasian blue tit LSPE   LC
Cyanopica cooki Iberian magpie LSPE   LC
Dendrocopos major Great spotted woodpecker LSPE   LC
Emberiza calandra Corn bunting LSPE   LC
Emberiza cia Rock bunting LSPE   NT
Emberiza cirlus Cirl bunting LSPE   NT
Emberiza hortulana Ortolan bunting LSPE   VU
Erithacus rubecula European robin LSPE   LC
Falco peregrinus Peregrine falcon LSPE Vulnerable NT
Falco tinnunculus Common kestrel LSPE   VU
Ficedula hypoleuca Pied flycatcher LSPE   LC
Fringilla coelebs Common chaffinch LSPE*   LC
Galerida theklae Thekla’s lark LSPE   LC
Garrulus glandarius Eurasian jay LSPE   LC
Gyps fulvus Griffon vulture LSPE   LC
Hieraetus pennatus Booted eagle LSPE Special Interest LC
Hippolais polyglotta Melodious warbler LSPE   LC
Hirundo rustica Barn swallow LSPE   LC
Lanius collurio Red-backed shrike LSPE   VU
Lanius meridionalis Great grey shrike LSPE Special Interest VU
Lanius senator Woodchat shrike LSPE   VU
Linaria cannabina Common linnet LSPE   LC
Lophophanes cristatus Crested tit LSPE   LC
Loxia curvirostra Red crossbill LSPE   LC
Lullula arborea Woodlark LSPE   VU
Luscinia megarhynchos Common nightingale LSPE   LC
Luscinia svecica Bluethroat LSPE Special Interest NT
Milvus migrans Black kite LSPE   LC
Milvus milvus Red kite EN Vulnerable EN
Monticola saxatilis Common rock thrush LSPE   VU
Monticola solitarius Blue rock thrush LSPE   LC
Motacilla alba White wagtail LSPE   LC
Motacilla cinerea Grey wagtail LSPE   LC
Motacilla flava Western yellow wagtail LSPE   LC
Oenanthe oenanthe Northern wheatear LSPE   VU
Oriolus oriolus Golden oriole LSPE   LC
Parus major Great tit LSPE   LC
Periparus ater Coal tit LSPE   LC
Pernis apivorus European honey-buzzard LSPE Special Interest NT
Petronia petronia Rock sparrow LSPE   LC
Phoenicurus ochruros Black redstart LSPE   LC
Phylloscopus bonelli Western Bonelli’s warbler LSPE   LC
Phylloscopus collybita Common chiffchaff LSPE   LC
Pica pica Eurasian magpie LSPE   LC
Picus sharpei Iberian green woodpecker LSPE   LC
Prunella modularis Dunnock LSPE   LC
Ptyonoprogne rupestris Crag martin LSPE   LC
Pyrrhocorax pyrrhocorax Red-billed chough LSPE   LC
Regulus ignicapilla Firecrest LSPE   LC
Regulus regulus Goldcrest LSPE   LC
Saxicola rubicola European stonechat LSPE   LC
Serinus serinus European serin LSPE   LC
Sitta europaea Eurasian nuthatch LSPE   LC
Streptopelia turtur European turtle dove LSPE   VU
Sturnus unicolor Spotless starling LSPE   LC
Sylvia atricapilla Eurasian blackcap LSPE   LC
Sylvia borin Garden warbler LSPE   LC
Troglodytes troglodytes Eurasian wren LSPE   LC
Turdus merula Common blackbird LSPE   LC
Turdus philomelos Song thrush LSPE   LC
Turdus viscivorus Mistle thrush LSPE   LC
Upupa epops Eurasian hoopoe LSPE   LC

Annex 2

List of the most abundant bird species present in the National Park according to the number of grid cells they occupy. Each species is indexed in order, indicating the percentage of 1×1 km grid cells in which it has been recorded relative to the total.

Scientific name English name % occupied grid cells
Fringilla coelebs Common chaffinch 66.36%
Periparus ater Coal tit 53.94%
Erithacus rubecula European robin 51.48%
Emberiza cia Rock bunting 44.55%
Prunella modularis Dunnock 41.82%
Troglodytes troglodytes Eurasian wren 38.79%
Lophophanes cristatus Crested tit 37.88%
Certhia brachydactyla Short-toed treecreeper 35.76%
Sylvia atricapilla Eurasian blackcap 35.45%
Regulus ignicapilla Firecrest 33.64%
Lullula arborea Woodlark 27.58%
Gyps fulvus Griffon vulture 24.85%
Turdus viscivorus Mistle thrush 24.55%
Turdus merula Common blackbird 24.24%
Alauda arvensis Skylark 23.33%
Saxicola rubicola European stonechat 22.73%
Columba palumbus Common wood pigeon 22.12%
Phylloscopus bonelli Western Bonelli’s warbler 20.61%
Garrulus glandarius Eurasian jay 18.18%
Cuculus canorus Common cuckoo 16.97%
Aegypius monachus Cinereous vulture 16.67%
Oenanthe oenanthe Northern wheatear 16.06%
Corvus corax Common raven 15.45%
Parus major Great tit 15.15%
Dendrocopos major Great spotted woodpecker 13.94%
Regulus regulus Goldcrest 13.33%
Linaria cannabina Eurasian linnet 12.73%
Emberiza hortulana Ortolan bunting 11.82%
Corvus corone Carrion crow 11.52%
Sitta europaea Eurasian nuthatch 11.52%
Phoenicurus ochruros Black redstart 11.52%
Loxia curvirostra Red crossbill 10.30%
Cyanistes caeruleus Eurasian blue tit 10.30%
Serinus serinus European serin 10.00%
Curruca iberiae Western subalpine warbler 9.70%
Aegithalos caudatus Long-tailed tit 9.70%
Curruca hortensis Western Orphean warbler 8.79%
Anthus campestris Tawny pipit 8.48%
Hieraaetus pennatus Booted eagle 7.58%
Curruca undata Dartford warbler 6.06%
Monticola saxatilis Rufous-tailed rock thrush 5.76%
Anthus spinoletta Water pipit 4.55%
Luscinia megarhynchos Common nightingale 4.55%
Sylvia borin Garden warbler 4.24%
Ficedula hypoleuca Pied flycatcher 4.24%
Alectoris rufa Red-legged partridge 4.24%
Picus sharpei Iberian green woodpecker 4.24%
Pyrrhocorax pyrrhocorax Red-billed chough 4.24%
Carduelis citrinella Citril finch 3.94%
Buteo buteo Common buzzard 3.94%
Monticola solitarius Blue rock thrush 3.94%
Apus apus Common swift 3.94%
Milvus milvus Red kite 3.64%
Luscinia svecica Bluethroat 3.64%
Coturnix coturnix Common quail 3.03%
Ptyonoprogne rupestris Eurasian crag martin 2.73%
Anthus trivialis Tree pipit 2.73%
Curruca melanocephala Sardinian warbler 2.73%
Hippolais polyglotta Melodious warbler 2.73%
Motacilla cinerea Grey wagtail 2.73%
Curruca conspicillata Spectacled warbler 2.42%
Emberiza cirlus Cirl bunting 2.42%
Lanius collurio Red-backed shrike 2.12%
Cyanopica cooki Iberian magpie 1.82%
Turdus philomelos Song thrush 1.82%
Sturnus unicolor Spotless starling 1.52%
Caprimulgus europaeus European nightjar 1.52%
Cecropis daurica Red-rumped swallow 1.52%
Accipiter gentilis Northern goshawk 1.21%
Accipiter nisus Eurasian sparrowhawk 1.21%
Emberiza calandra Corn bunting 0.91%
Chloris chloris European greenfinch 0.91%
Petronia petronia Rock sparrow 0.91%
Falco tinnunculus Common kestrel 0.91%
Motacilla flava Yellow wagtail 0.91%
Oriolus oriolus Eurasian golden oriole 0.30%
Hirundo daurica Red-rumped swallow 0.30%
Pernis apivorus European honey buzzard 0.30%
Falco peregrinus Peregrine falcon 0.30%
Aquila fasciata Bonelli’s eagle 0.30%
Cocothraustes cocothraustes Hawfinch 0.30%
Muscicapa striata Spotted flycatcher 0.30%
Aquila adalberti Spanish imperial eagle 0.30%
Cinclus cinclus White-throated dipper 0.30%
Pica pica Eurasian magpie 0.30%
Streptopelia turtur European turtle dove 0.30%