CR – Critically Endangered, EN – Endangered, VU – Vulnerable, DD – Data Deficient, NT – Near Threatened
The fieldwork was carried out across the 21,714 hectares occupied by the Sierra de Guadarrama National Park within the Community of Madrid. Of the 330 grid squares that include part of their surface inside the park, surveys were conducted in 282 of them in order to determine the complete species list and the relative abundance of birds in each one. The 48 grid squares that were not visited correspond to those with a very small area inside the National Park. Within each surveyed square, all environments or habitats present were explored, since each one may host different species.
This national park lies at a considerable altitude above sea level (900–2,490 m), which strongly influences the types of habitat found there. Alpine grasslands, humid meadows, and mountain grasslands predominate, but there are also large forested areas. Pine and oak forests are common, and holm oak stands appear to a lesser extent. Shrubland is also an important component, especially dominated by broom and heath.
A total of 98 species were recorded, detailed in Annex 1. All of them are breeding species in the area. The distribution map and the habitats used by each species are included in the section Species Index.
The most widely distributed or most common species, according to the number of grid squares and in order of importance, are: common chaffinch, coal tit, European robin, rock bunting, dunnock, Eurasian wren, crested tit, and Eurasian blackcap. These eight species are present in more than 35% of the grid squares within the study area. In addition to these, three major groups can be identified:
In addition to the species recorded with signs of breeding, other species were also identified that are considered migratory or incidental observations, given the mobility of this group of fauna, which can result in sightings of dispersed individuals of almost any species anywhere. These include: common house martin, common redstart, willow warbler, greater short-toed lark, Eurasian collared dove, and brambling.
Species inventory by UTM 1×1 km grid
The number of species present in each grid square represents the specific richness per area in the park, or the species richness of each individual square. The 1×1 km grid squares shown in dark colours in Figure 1 represent the areas with the highest species richness in the national park.
The grid squares with the highest number of species (more than 20 recorded species) are distributed across different zones of the park, following patches of vegetation with greater diversity and more varied environments. Areas with lower species richness correspond to higher altitudes with little or no vegetation. Notable areas with higher diversity include the oak forests on the slopes extending above the village of Rascafría and the pine forests on both sides of the Navafría pass. Higher-richness areas also occur on the Morcuera slope towards Rascafría, the area known as El Pinganillo, and especially Las Hoyuelas (Figures 1, 2, and 3). Equally notable are the species-rich areas of the Hueco de San Blas and the southern slopes of Morcuera. The species found in these forested areas constitute typical mountain forest communities, including birds native to the predominant pine forest, as well as species from mixed forests, oak woods, and shrublands where these formations occur.
Most of the grid squares with a low number of species (1 to 5), and a large proportion of those with few species (6 to 10) are located mainly in the highest areas, that is, the peaks of the National Park, and in rocky slopes with little vegetation. These areas correspond to zones dominated by rocky habitats, high mountain grasslands, and broom scrub. In these habitats, species richness and bird abundance decline markedly, although broom scrub areas can host high densities of birds during the breeding season, with skylarks, dunnocks, and European stonechats being the main representatives.
A group of species consistently present within the park boundaries are the birds of prey. The colonies of cinereous vulture in Rascafría and La Pedriza, along with the small colonies of griffon vulture distributed across various rocky areas of the mountains, contribute a significant number of individuals, and their ease of detection means that visitors are almost constantly observing large birds in the park. Other birds of prey in higher threat categories, such as the Spanish imperial eagle or Bonelli’s eagle, are represented much more sparsely.
The presence of rare and striking species such as the common rock thrush and the bluethroat is also notable. Some species that used to be more abundant, although detected during this atlas’ fieldwork, are now in decline and scarcely represented; this is the case of the alpine accentor.



Threatened species and their distribution
Las especies de fauna están catalogadas de distinta manera en los diferentes catálogos de especies protegidas según la escala (estatal o regional) o según criterios específicos estatales o internacionales (Acuerdo del Consejo de Ministros de 24 de febrero de 2017, por el que se aprueban los criterios orientadores para la inclusión de taxones y poblaciones en el Catálogo Español de Especies Amenazadas o UICN).
Para que quede debidamente reflejada la distribución de las especies según estos criterios, a continuación se incluyen los listados y la distribución de las especies según su grado de amenaza considerando los siguientes documentos: Catálogo Español de Especies Amenazadas y Listado de Especies Silvestres en Régimen de Protección Especial, Libro Rojo de las Aves de España, Catálogo Regional de especies amenazadas de fauna y flora silvestres de la Comunidad de Madrid.
Threatened species according to the Spanish Catalogue of Threatened Species
Of the 98 species present in the Park during the breeding season, 78 are included in the List of Wild Species under Special Protection, and 4 of them appear in the Spanish Catalogue of Threatened Species with a very unfavourable conservation status: Endangered (2 species) and Vulnerable (2 species; tables 1, 2 and 3). The full list of species and their category is included in Annex 2.
| Category | Number of species |
|---|---|
| Endangered | 2 |
| Vulnerable | 2 |
| Included in the List of Wild Species under Special Protection | 78 |
Table 1. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the Spanish Catalogue of Threatened Species in Spain.
Endangered
Spanish imperial eagle
Red kite
Table 2. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Spanish Catalogue of Threatened Species in Spain.
Vulnerable
Cinereous vulture
Bonelli’s eagle
Table 3. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Vulnerable according to the Spanish Catalogue of Threatened Species in Spain.
Considering the Spanish Catalogue of Threatened Species, the following areas would be identified as occupied by species in the highest threat categories within that catalogue:

Figure 4. Areas with the presence of species classified as Endangered in the Spanish Catalogue of Threatened Species.

Figure 5. Areas with the presence of species included in the Vulnerable category in the Spanish Catalogue of Threatened Species.
Threatened species according to the Red Book of the Birds of Spain
Of the 98 species present in the National Park during the breeding season, 14 are in an unfavourable conservation status and are included in the threat categories of Endangered (7 species) and Vulnerable (7 species) in the latest Red List prepared by SEO/BirdLife for the Ministry for the Ecological Transition and the Demographic Challenge (tables 4, 5 and 6).
| Category | Number of species |
|---|---|
| Endangered | 7 |
| Vulnerable | 7 |
| Near Threatened | 13 |
| Least Concern | 70 |
| Data Deficient | 2 |
Table 4. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).
Endangered
Spanish imperial eagle
Woodchat shrike
Great grey shrike
Common kestrel
Common quail
Dartford warbler
Red kite
Table 5. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).
Vulnerable
Bonelli’s eagle
Red-backed shrike
Skylark
Barn swallow
Red-legged partridge
European turtle dove
Common swift
Table 6. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Vulnerable in the latest Red Book and Red List of Birds in Spain (SEO/BirdLife, 2021).
The distribution of species included in any unfavourable conservation status must guide the management of a protected area. In this case, the species detected as breeders within the boundaries considered in this study, and included in two of the highest categories according to IUCN criteria, highlight the priority conservation areas in this regard (figures 6 and 7).

Figure 6. Grid squares with the presence of species classified as Endangered in the latest Red Book.

Figure 7. Grid squares with the presence of species included in the Vulnerable category in the latest Red Book.
Threatened species according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid
Of the 98 species present in the National Park during the breeding season, 17 are in an unfavourable conservation status and are included in the threat categories of Endangered (3 species), Vulnerable (2 species), Special Interest (1 species), and Sensitive to Habitat Alteration (11 species) in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid (tables 7 and 8).
| Category | Number of species |
|---|---|
| Endangered | 3 |
| Vulnerable | 2 |
| Special Interest | 1 |
| Sensitive to Habitat Alteration | 11 |
Table 7. Number of species present during the breeding season in the Sierra de Guadarrama National Park according to their threat category in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.
Endangered
Spanish imperial eagle
Bonelli’s eagle
Cinereous vulture
Table 8. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.
Considering the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid, the following areas would be identified as occupied by species in the highest categories of that catalogue:

Figure 8. Number of species present during the breeding season in the Sierra de Guadarrama National Park classified as Endangered according to the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.

Figure 9. Grid squares with the presence of species included in the Vulnerable category in the Regional Catalogue of Wild Fauna and Flora of the Community of Madrid.
Spanish Catalogue of Threatened Species and List of Wild Species under Special Protection:
EN – Endangered, VU – Vulnerable, LSPE – Included in the List of Wild Species under Special Protection.
Red Book of the Birds of Spain:
CR – Critically Endangered, EN – Endangered, VU – Vulnerable, DD – Data Deficient, NT – Near Threatened
Regional Catalogue of Threatened Species of Wild Fauna and Flora:
EN – Endangered, SAH – Sensitive to Habitat Alteration, VU – Vulnerable, DIE – Special Interest.
Annex 1
List of bird species present in the National Park. It is indexed by species name for quick identification and consultation of a specific species.
| Scientific name | Common name (Spanish) | CEA 2024 | Madrid 1992 | Red Book 2021 |
|---|---|---|---|---|
| Accipiter gentilis | Northern goshawk | LSPE | LC | |
| Accipiter nisus | Eurasian sparrowhawk | LSPE | LC | |
| Aegithalos caudatus | Long-tailed tit | LSPE | LC | |
| Aegypius monachus | Cinereous vulture | VU | Endangered | NT |
| Alauda arvensis | Skylark | LSPE | VU | |
| Alectoris rufa | Red-legged partridge | LSPE | VU | |
| Anthus campestris | Tawny pipit | LSPE | VU | |
| Anthus spinoletta | Water pipit | LSPE | LC | |
| Anthus trivialis | Tree pipit | LSPE | LC | |
| Apus apus | Common swift | LSPE | LC | |
| Aquila adalberti | Spanish imperial eagle | EN | Endangered | VU |
| Aquila chrysaetos | Golden eagle | LSPE | Sensitive to Habitat Alteration | LC |
| Aquila fasciata | Bonelli’s eagle | VU | Endangered | VU |
| Buteo buteo | Common buzzard | LSPE | LC | |
| Caprimulgus europaeus | European nightjar | LSPE | LC | |
| Carduelis carduelis | European goldfinch | LSPE | LC | |
| Carduelis citrinella | Citril finch | LSPE | Special Interest | NT |
| Cecropis daurica | Red-rumped swallow | LSPE | LC | |
| Certhia brachydactyla | Short-toed treecreeper | LSPE | LC | |
| Chloris chloris | European greenfinch | LSPE | LC | |
| Cinclus cinclus | White-throated dipper | LSPE | Special Interest | LC |
| Circataetus gallicus | Short-toed snake eagle | LSPE | LC | |
| Cocothraustes cocothraustes | Hawfinch | LSPE | Special Interest | LC |
| Columba palumbus | Common wood pigeon | LSPE | LC | |
| Corvus corax | Common raven | LC | ||
| Corvus corone | Carrion crow | LC | ||
| Coturnix coturnix | Common quail | LSPE | LC | |
| Cuculus canorus | Common cuckoo | LSPE | LC | |
| Curruca communis | Common whitethroat | LSPE | LC | |
| Curruca conspicillata | Spectacled warbler | LSPE | LC | |
| Curruca hortensis | Western Orphean warbler | LSPE | Special Interest | LC |
| Curruca iberiae | Western subalpine warbler | LSPE | LC | |
| Curruca melanocephala | Sardinian warbler | LSPE | LC | |
| Curruca undata | Dartford warbler | LSPE | VU | |
| Cyanistes caeruleus | Eurasian blue tit | LSPE | LC | |
| Cyanopica cooki | Iberian magpie | LSPE | LC | |
| Dendrocopos major | Great spotted woodpecker | LSPE | LC | |
| Emberiza calandra | Corn bunting | LSPE | LC | |
| Emberiza cia | Rock bunting | LSPE | NT | |
| Emberiza cirlus | Cirl bunting | LSPE | NT | |
| Emberiza hortulana | Ortolan bunting | LSPE | VU | |
| Erithacus rubecula | European robin | LSPE | LC | |
| Falco peregrinus | Peregrine falcon | LSPE | Vulnerable | NT |
| Falco tinnunculus | Common kestrel | LSPE | VU | |
| Ficedula hypoleuca | Pied flycatcher | LSPE | LC | |
| Fringilla coelebs | Common chaffinch | LSPE* | LC | |
| Galerida theklae | Thekla’s lark | LSPE | LC | |
| Garrulus glandarius | Eurasian jay | LSPE | LC | |
| Gyps fulvus | Griffon vulture | LSPE | LC | |
| Hieraetus pennatus | Booted eagle | LSPE | Special Interest | LC |
| Hippolais polyglotta | Melodious warbler | LSPE | LC | |
| Hirundo rustica | Barn swallow | LSPE | LC | |
| Lanius collurio | Red-backed shrike | LSPE | VU | |
| Lanius meridionalis | Great grey shrike | LSPE | Special Interest | VU |
| Lanius senator | Woodchat shrike | LSPE | VU | |
| Linaria cannabina | Common linnet | LSPE | LC | |
| Lophophanes cristatus | Crested tit | LSPE | LC | |
| Loxia curvirostra | Red crossbill | LSPE | LC | |
| Lullula arborea | Woodlark | LSPE | VU | |
| Luscinia megarhynchos | Common nightingale | LSPE | LC | |
| Luscinia svecica | Bluethroat | LSPE | Special Interest | NT |
| Milvus migrans | Black kite | LSPE | LC | |
| Milvus milvus | Red kite | EN | Vulnerable | EN |
| Monticola saxatilis | Common rock thrush | LSPE | VU | |
| Monticola solitarius | Blue rock thrush | LSPE | LC | |
| Motacilla alba | White wagtail | LSPE | LC | |
| Motacilla cinerea | Grey wagtail | LSPE | LC | |
| Motacilla flava | Western yellow wagtail | LSPE | LC | |
| Oenanthe oenanthe | Northern wheatear | LSPE | VU | |
| Oriolus oriolus | Golden oriole | LSPE | LC | |
| Parus major | Great tit | LSPE | LC | |
| Periparus ater | Coal tit | LSPE | LC | |
| Pernis apivorus | European honey-buzzard | LSPE | Special Interest | NT |
| Petronia petronia | Rock sparrow | LSPE | LC | |
| Phoenicurus ochruros | Black redstart | LSPE | LC | |
| Phylloscopus bonelli | Western Bonelli’s warbler | LSPE | LC | |
| Phylloscopus collybita | Common chiffchaff | LSPE | LC | |
| Pica pica | Eurasian magpie | LSPE | LC | |
| Picus sharpei | Iberian green woodpecker | LSPE | LC | |
| Prunella modularis | Dunnock | LSPE | LC | |
| Ptyonoprogne rupestris | Crag martin | LSPE | LC | |
| Pyrrhocorax pyrrhocorax | Red-billed chough | LSPE | LC | |
| Regulus ignicapilla | Firecrest | LSPE | LC | |
| Regulus regulus | Goldcrest | LSPE | LC | |
| Saxicola rubicola | European stonechat | LSPE | LC | |
| Serinus serinus | European serin | LSPE | LC | |
| Sitta europaea | Eurasian nuthatch | LSPE | LC | |
| Streptopelia turtur | European turtle dove | LSPE | VU | |
| Sturnus unicolor | Spotless starling | LSPE | LC | |
| Sylvia atricapilla | Eurasian blackcap | LSPE | LC | |
| Sylvia borin | Garden warbler | LSPE | LC | |
| Troglodytes troglodytes | Eurasian wren | LSPE | LC | |
| Turdus merula | Common blackbird | LSPE | LC | |
| Turdus philomelos | Song thrush | LSPE | LC | |
| Turdus viscivorus | Mistle thrush | LSPE | LC | |
| Upupa epops | Eurasian hoopoe | LSPE | LC |
Annex 2
List of the most abundant bird species present in the National Park according to the number of grid cells they occupy. Each species is indexed in order, indicating the percentage of 1×1 km grid cells in which it has been recorded relative to the total.
| Scientific name | English name | % occupied grid cells |
|---|---|---|
| Fringilla coelebs | Common chaffinch | 66.36% |
| Periparus ater | Coal tit | 53.94% |
| Erithacus rubecula | European robin | 51.48% |
| Emberiza cia | Rock bunting | 44.55% |
| Prunella modularis | Dunnock | 41.82% |
| Troglodytes troglodytes | Eurasian wren | 38.79% |
| Lophophanes cristatus | Crested tit | 37.88% |
| Certhia brachydactyla | Short-toed treecreeper | 35.76% |
| Sylvia atricapilla | Eurasian blackcap | 35.45% |
| Regulus ignicapilla | Firecrest | 33.64% |
| Lullula arborea | Woodlark | 27.58% |
| Gyps fulvus | Griffon vulture | 24.85% |
| Turdus viscivorus | Mistle thrush | 24.55% |
| Turdus merula | Common blackbird | 24.24% |
| Alauda arvensis | Skylark | 23.33% |
| Saxicola rubicola | European stonechat | 22.73% |
| Columba palumbus | Common wood pigeon | 22.12% |
| Phylloscopus bonelli | Western Bonelli’s warbler | 20.61% |
| Garrulus glandarius | Eurasian jay | 18.18% |
| Cuculus canorus | Common cuckoo | 16.97% |
| Aegypius monachus | Cinereous vulture | 16.67% |
| Oenanthe oenanthe | Northern wheatear | 16.06% |
| Corvus corax | Common raven | 15.45% |
| Parus major | Great tit | 15.15% |
| Dendrocopos major | Great spotted woodpecker | 13.94% |
| Regulus regulus | Goldcrest | 13.33% |
| Linaria cannabina | Eurasian linnet | 12.73% |
| Emberiza hortulana | Ortolan bunting | 11.82% |
| Corvus corone | Carrion crow | 11.52% |
| Sitta europaea | Eurasian nuthatch | 11.52% |
| Phoenicurus ochruros | Black redstart | 11.52% |
| Loxia curvirostra | Red crossbill | 10.30% |
| Cyanistes caeruleus | Eurasian blue tit | 10.30% |
| Serinus serinus | European serin | 10.00% |
| Curruca iberiae | Western subalpine warbler | 9.70% |
| Aegithalos caudatus | Long-tailed tit | 9.70% |
| Curruca hortensis | Western Orphean warbler | 8.79% |
| Anthus campestris | Tawny pipit | 8.48% |
| Hieraaetus pennatus | Booted eagle | 7.58% |
| Curruca undata | Dartford warbler | 6.06% |
| Monticola saxatilis | Rufous-tailed rock thrush | 5.76% |
| Anthus spinoletta | Water pipit | 4.55% |
| Luscinia megarhynchos | Common nightingale | 4.55% |
| Sylvia borin | Garden warbler | 4.24% |
| Ficedula hypoleuca | Pied flycatcher | 4.24% |
| Alectoris rufa | Red-legged partridge | 4.24% |
| Picus sharpei | Iberian green woodpecker | 4.24% |
| Pyrrhocorax pyrrhocorax | Red-billed chough | 4.24% |
| Carduelis citrinella | Citril finch | 3.94% |
| Buteo buteo | Common buzzard | 3.94% |
| Monticola solitarius | Blue rock thrush | 3.94% |
| Apus apus | Common swift | 3.94% |
| Milvus milvus | Red kite | 3.64% |
| Luscinia svecica | Bluethroat | 3.64% |
| Coturnix coturnix | Common quail | 3.03% |
| Ptyonoprogne rupestris | Eurasian crag martin | 2.73% |
| Anthus trivialis | Tree pipit | 2.73% |
| Curruca melanocephala | Sardinian warbler | 2.73% |
| Hippolais polyglotta | Melodious warbler | 2.73% |
| Motacilla cinerea | Grey wagtail | 2.73% |
| Curruca conspicillata | Spectacled warbler | 2.42% |
| Emberiza cirlus | Cirl bunting | 2.42% |
| Lanius collurio | Red-backed shrike | 2.12% |
| Cyanopica cooki | Iberian magpie | 1.82% |
| Turdus philomelos | Song thrush | 1.82% |
| Sturnus unicolor | Spotless starling | 1.52% |
| Caprimulgus europaeus | European nightjar | 1.52% |
| Cecropis daurica | Red-rumped swallow | 1.52% |
| Accipiter gentilis | Northern goshawk | 1.21% |
| Accipiter nisus | Eurasian sparrowhawk | 1.21% |
| Emberiza calandra | Corn bunting | 0.91% |
| Chloris chloris | European greenfinch | 0.91% |
| Petronia petronia | Rock sparrow | 0.91% |
| Falco tinnunculus | Common kestrel | 0.91% |
| Motacilla flava | Yellow wagtail | 0.91% |
| Oriolus oriolus | Eurasian golden oriole | 0.30% |
| Hirundo daurica | Red-rumped swallow | 0.30% |
| Pernis apivorus | European honey buzzard | 0.30% |
| Falco peregrinus | Peregrine falcon | 0.30% |
| Aquila fasciata | Bonelli’s eagle | 0.30% |
| Cocothraustes cocothraustes | Hawfinch | 0.30% |
| Muscicapa striata | Spotted flycatcher | 0.30% |
| Aquila adalberti | Spanish imperial eagle | 0.30% |
| Cinclus cinclus | White-throated dipper | 0.30% |
| Pica pica | Eurasian magpie | 0.30% |
| Streptopelia turtur | European turtle dove | 0.30% |